1.Herbs perennial. Rhizomes black-brown, usually torulose, large.
多年生草本根状茎黑棕色，通常，大。
2.valves with a prominent midvein and often distinct marginal veins, glabrous or glandular, torulose;
裂爿具一突出中脉和通常离生边缘脉，无毛或具腺，；
3.valves papery, with a prominent midvein and distinct lateral veins, glabrous, torulose;
裂爿纸质，具一突出和离生侧脉，无毛，近念珠状；
4.valvular segment dehiscent, few to many seeded, longer or shorter than terminal segment, torulose;
瓣裂裂片开裂，少数到多数种子，长或短于顶生裂片，；
5.valves with a prominent midvein, smooth or torulose;
裂爿具一突出中脉，滤波或；